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        <gco:CharacterString>Dillwyn’s Dung Beetle Onthophagus nuchicornis was formerly local but widespread across England and Wales. It has declined dramatically in England and since 2000 is extant at just five of the forty known locations. In Wales, this species has fared better but is restricted to grazed coastal sand dune systems. Historically known from fifteen sites, it was still present at half of these in 2017. The beetle feeds on the dung of a range of mammals but is probably dependent for breeding on medium to large herbivore dung such as cattle, ponies and sheep.  A survey in 2017 of all the known sites in Wales found Onthophagus nuchicornis at seven localities – Kenfig Burrows (975 adults), Merthyr Mawr Warren (36), Morfa Harlech (1), Oxwich Burrows (4), Pembrey Burrows (32), Pennard Burrows (4) and Whiteford Burrows (5). The authors regard the populations at Kenfig Burrows and Methyr Mawr to be the best in the UK. Elsewhere, absence and low population levels are strongly correlated to the cessation of grazing and the quantity and quality of available dung on sand.  Following the 2017 survey, a more intensive survey of Morfa Harlech, which represents a northern outlier for the beetle, was undertaken in 2019 to determine the status of Onthophagus nuchicornis at this site.</gco:CharacterString>
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